Today is "where did that idiom come from" day. Used to express disbelief or surprise and amazement,the phrase "I'll be a monkey's uncle" is said to date from 1925 and the widely publicized Scopes Trial. It was a sarcastic remark made by people opposed to the theory of evolution. Darwin's book "The Origin of Species" was published in 1859,followed by "The Descent of Man" in 1871,the idea of humans being descendants of apes was considered blasphemous,so the phrase was put into use to express disbelief in Darwin's theory.
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